So karst vegetation is charactered by petrophile , xerophile and calciphihe in those areas where soil cover is very thin , and the underground drainage system is well developed . the running of karst ecosystem is controlled by two subsystems , which are karst dynamic system in inorganic life and genetic information transmitting system in organic life 岩溶生态系统的运行受到两个子系统的联合作用驱动,即无机方面的“岩溶动力系统”和生命方面的“遗传信息传递系统” ,但主要受以碳、水、钙循环为中心的岩溶动力系统的控制。