Embryological studies reveal that megaspore and such cells as egg , synergid and antipodals in mature embryo sac initiate the division of forming haploid plants through embryogenesis or callus formation 胚胎学观察揭示大孢子与胚囊内的卵细胞、助细胞和反足细胞均有可能在培养中启动分裂,通过胚状体或愈伤组织形成单倍体植株。